Taxonomic Group	Name	Year	Invasion Status	Population Status	Vectors
Fishes	Alosa sapidissima	1880	Definite	Established	Fisheries Accidental (not Oyster)
Fishes	Ameiurus catus	1930	Definite	Established	Fisheries Intentional
Fishes	Ameiurus melas	1894	Definite	Established	Fisheries Intentional
Fishes	Ameiurus natalis	1905	Definite	Established	Fisheries Intentional
Fishes	Ameiurus nebulosus	1894	Definite	Established	Fisheries Intentional
Crustaceans-Barnacles	Amphibalanus improvisus	1957	Definite	Established	Ballast Water, Hull Fouling
Crustaceans-Cladocerans	Bosmina coregoni	2007	Definite	Established	Aquatic Plant Shipments, Ballast Water, Fisheries Accidental (not Oyster), Natural Dispersal
Crustaceans-Isopods	Caecidotea racovitzai	1999	Definite	Established	Aquatic Plant Shipments, Ballast Water
Fishes	Carassius auratus	1933	Definite	Established	None
Mollusks-Gastropods	Cipangopaludina chinensis	2002	Definite	Established	Aquatic Plant Shipments, Fisheries Accidental (not Oyster), Fisheries Intentional, Pet Release
Bryozoans	Conopeum chesapeakensis	2015	Definite	Established	Ballast Water, Hull Fouling
Mollusks-Bivalves	Corbicula fluminea	1938	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster), Fisheries Intentional
Cnidarians-Hydrozoans	Cordylophora caspia	1965	Definite	Established	Aquatic Plant Shipments, Fisheries Accidental (not Oyster), Hull Fouling, Oyster Accidental
Crustaceans-Amphipods	Crangonyx pseudogracilis	2002	Definite	Established	Aquatic Plant Shipments, Ballast Water, Fisheries Accidental (not Oyster)
Fishes	Cyprinus carpio	1883	Definite	Established	Fisheries Accidental (not Oyster), Fisheries Intentional
Annelids-Oligochaetes	Eukerria saltensis*	2003	Definite	Established	Agricultural Weed, Aquatic Plant Shipments, Cargo
Bryozoans	Fredericella indica	1999	Definite	Established	Aquatic Plant Shipments, Fisheries Accidental (not Oyster)
Fishes	Fundulus diaphanus*	1971	Definite	Established	Discarded Bait, Fisheries Accidental (not Oyster)
Arthropoda-Insects	Galerucella calmariensis*	1997	Definite	Established	Biocontrol
Arthropoda-Insects	Galerucella pusilla*	1997	Definite	Established	Biocontrol
Fishes	Gambusia affinis	1995	Definite	Established	None
Crustaceans-Amphipods	Grandidierella japonica	1999	Definite	Established	Ballast Water, Hull Fouling, Natural Dispersal
Annelids-Polychaetes	Hediste diadroma	2009	Definite	Established	Ballast Water
Annelids-Polychaetes	Heteromastus filiformis species complex	2002	Definite	Established	Ballast Water, Hull Fouling
Annelids-Polychaetes	Hobsonia florida	1979	Definite	Established	Ballast Water
Arthropoda-Insects	Hylobius transversovittatus*	1997	Definite	Established	Biocontrol
Fishes	Ictalurus punctatus	1945	Definite	Established	None
Fishes	Lepomis gibbosus	1893	Definite	Established	Fisheries Intentional
Fishes	Lepomis gulosus	1930	Definite	Established	Fisheries Intentional
Fishes	Lepomis macrochirus	1933	Definite	Established	Fisheries Accidental (not Oyster)
Crustaceans-Copepods	Lernaea cyprinacea	1948	Definite	Established	Fisheries Accidental (not Oyster), Pet Release
Crustaceans-Copepods	Limnoithona tetraspina	2003	Definite	Established	Ballast Water
Annelids-Polychaetes	Marenzelleria neglecta	2020	Definite	Established	Biocontrol
Fishes	Micropterus dolomieu	1924	Definite	Established	Fisheries Intentional
Fishes	Micropterus salmoides	1888	Definite	Established	None
Fishes	Misgurnus anguillicaudatus*	1997	Definite	Established	None
Fishes	Morone saxatilis	1936	Definite	Established	Natural Dispersal
Mollusks-Bivalves	Mya arenaria	1973	Definite	Established	Fisheries Intentional
Arthropoda-Insects	Nanophyes marmoratus*	1997	Definite	Established	Biocontrol
Cnidarians-Anthozoans	Nematostella vectensis	1994	Definite	Established	Ballast Water, Hull Fouling
Crustaceans-Cumaceans	Nippoleucon hinumensis	1994	Definite	Established	Ballast Water
Fishes	Notemigonus crysoleucas	2002	Definite	Established	Discarded Bait
Mollusks-Bivalves	Nuttallia obscurata	2003	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster), Natural Dispersal
Crustaceans-Shrimp	Palaemon modestus	1995	Definite	Established	Ballast Water
Bryozoans	Pectinatella magnifica	1998	Definite	Established	Aquatic Plant Shipments, Fisheries Accidental (not Oyster)
Fishes	Perca flavescens	1891	Definite	Established	Fisheries Accidental (not Oyster)
Annelids-Polychaetes	Polydora cornuta	1980	Definite	Established	Ballast Water, Hull Fouling
Fishes	Pomoxis annularis	1893	Definite	Established	Fisheries Intentional
Fishes	Pomoxis nigromaculatus	1893	Definite	Established	Fisheries Intentional
Mollusks-Gastropods	Potamopyrgus antipodarum	1997	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster)
Crustaceans-Copepods	Pseudodiaptomus forbesi	1999	Definite	Established	Ballast Water
Annelids-Polychaetes	Pseudopolydora cf. kempi	1980	Definite	Established	Ballast Water, Hull Fouling
Fishes	Rhinogobius brunneus species complex*	2004	Definite	Established	Ballast Water
Fishes	Sander vitreus*	1981	Definite	Established	Fisheries Intentional
Crustaceans-Tanaids	Sinelobus cf. stanfordi	2003	Definite	Established	Ballast Water, Hull Fouling
Crustaceans-Copepods	Sinocalanus doerrii	1999	Definite	Established	Ballast Water
Annelids-Polychaetes	Streblospio benedicti	1990	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster), Hull Fouling
Annelids-Oligochaetes	Stylodrilus heringianus*	2001	Definite	Established	Aquatic Plant Shipments, Ballast Water, Fisheries Accidental (not Oyster)
Entoprocts	Urnatella gracilis	2002	Definite	Established	Aquatic Plant Shipments, Fisheries Accidental (not Oyster)

* = Species summary still under review.

p260 (Columbia River) Summary