Taxonomic Group	Name	Year	Invasion Status	Population Status	Vectors
Cnidarians-Anthozoans	Actinia equina*	2021	Definite	Established	Ballast Water, Hull Fouling
Cnidarians-Anthozoans	Aiptasiogeton eruptaurantia*	2010	Definite	Established	Hull Fouling
Fishes	Ambloplites rupestris*	1842	Definite	Established	None
Fishes	Ameiurus catus	1949	Definite	Established	None
Fishes	Amia calva*	1942	Definite	Established	None
Crustaceans-Barnacles	Amphibalanus amphitrite	1967	Definite	Established	Ballast Water, Hull Fouling, Natural Dispersal
Nematodes	Anguillicola crassus*	1997	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster)
Arthropoda-Insects	Anisolabis maritima	1901	Definite	Established	Cargo, Dry Ballast
Algae	Antithamnion hubbsii*	1985	Definite	Established	Ballast Water, Hull Fouling
Fishes	Aplodinotus grunniens*	1978	Definite	Established	None
Crustaceans-Copepods	Argulus japonicus	1933	Definite	Established	None
Tunicates	Ascidiella aspersa	1983	Definite	Established	Hull Fouling
Entoprocts	Barentsia benedeni	1977	Definite	Established	Hull Fouling
Algae	Batophora oerstedii*	2015	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster), Hull Fouling
Mollusks-Gastropods	Bithynia tentaculata	1879	Definite	Established	None
Cnidarians-Hydrozoans	Blackfordia virginica	1904	Definite	Established	Ballast Water, Hull Fouling
Annelids-Polychaetes	Boccardiella ligerica	1990	Definite	Established	Ballast Water
Algae	Bonnemaisonia hamifera	1927	Definite	Established	Ballast Water, Hull Fouling
Tunicates	Botrylloides violaceus	1980	Definite	Established	Hull Fouling
Arthropoda-Insects	Brachydeutera longipes*	1983	Definite	Established	None
Annelids-Oligochaetes	Branchiura sowerbyi*	1955	Definite	Established	None
Algae	Bryopsis maxima*	2013	Definite	Established	Ballast Water, Hull Fouling
Bryozoans	Bugula neritina	1985	Definite	Established	Hull Fouling
Bryozoans	Bugulina simplex	1872	Definite	Established	Hull Fouling
Annelids-Oligochaetes	Cambarincola pamelae*	2003	Definite	Established	None
Crustaceans-Amphipods	Caprella mutica	1998	Definite	Established	Ballast Water, Hull Fouling
Fishes	Carassius auratus	1839	Definite	Established	None
Crustaceans-Crabs	Carcinus maenas	1817	Definite	Established	Dry Ballast, Hull Fouling
Algae	Ceramium secundatum*	2007	Definite	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Chaetococcus phragmitis*	2000	Definite	Established	None
Fishes	Channa argus	2003	Definite	Established	None
Crustaceans-Amphipods	Chelura terebrans	1875	Definite	Established	Hull Fouling
Arthropoda-Insects	Chilacis typhae*	1986	Definite	Established	None
Crustaceans-Barnacles	Chthamalus fragilis	1898	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Tunicates	Ciona intestinalis	1850	Definite	Established	Hull Fouling
Mollusks-Gastropods	Cipangopaludina chinensis	1925	Definite	Established	None
Tunicates	Clavelina lepadiformis	2009	Definite	Established	Hull Fouling
Algae	Codium fragile ssp. Fragile	1957	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Algae	Colpomenia peregrina*	2012	Definite	Established	Ballast Water, Hull Fouling
Platyhelminthes	Convoluta convoluta*	1995	Definite	Established	None
Mollusks-Bivalves	Corbicula fluminea	1971	Definite	Established	None
Cnidarians-Hydrozoans	Cordylophora caspia	1865	Definite	Established	Hull Fouling
Crustaceans-Amphipods	Corophium volutator	2001	Definite	Established	None
Cnidarians-Hydrozoans	Craspedacusta sowerbii	1884	Definite	Established	None
Platyhelminthes	Cryptocotyle lingua*	1911	Definite	Established	Dry Ballast, Fisheries Intentional, Hull Fouling, Natural Dispersal
Mollusks-Gastropods	Cuthona perca	1994	Definite	Established	Ballast Water, Hull Fouling
Algae	Cylindrospermopsis raciborskii*	1977	Definite	Established	None
Fishes	Cyprinus carpio	1832	Definite	Established	None
Mollusks-Bivalves	Cyrenoida floridana	1952	Definite	Established	Canal, Dry Ballast, Unknown Vector
Crustaceans-Cladocerans	Daphnia lumholtzi	1999	Definite	Established	None
Algae	Dasysiphonia japonica	2009	Definite	Established	Ballast Water, Hull Fouling
Cnidarians-Anthozoans	Diadumene lineata	1892	Definite	Established	Hull Fouling, Oyster Accidental
Tunicates	Didemnum vexillum	2000	Definite	Established	Hull Fouling
Tunicates	Diplosoma listerianum	1975	Definite	Established	Hull Fouling
Mollusks-Nudibranchs	Doris pseudoargus	2017	Definite	Established	None
Fishes	Dorosoma petenense	1953	Definite	Established	Fisheries Intentional
Mollusks-Bivalves	Dreissena bugensis	2008	Definite	Established	None
Mollusks-Bivalves	Dreissena polymorpha	1991	Definite	Established	None
Algae	Dumontia contorta*	1927	Definite	Established	Ballast Water, Hull Fouling
Tunicates	Ecteinascidia turbinata	1961	Definite	Established	Hull Fouling
Mollusks-Gastropods	Elimia livescens	1872	Definite	Established	None
Crustaceans-Crabs	Eriocheir sinensis	2007	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster), Fisheries Intentional
Fishes	Esox lucius*	1888	Definite	Established	None
Fishes	Etheostoma blennioides*	2011	Definite	Established	None
Fishes	Etheostoma zonale*	1984	Definite	Established	None
Crustaceans-Crayfish	Faxonius rusticus	1996	Definite	Established	None
Crustaceans-Crayfish	Faxonius virilis	1960	Definite	Established	None
Annelids-Polychaetes	Ficopomatus enigmaticus	1976	Definite	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Galerucella calmariensis*	1992	Definite	Established	None
Arthropoda-Insects	Galerucella pusilla*	1992	Definite	Established	None
Fishes	Gambusia affinis	1986	Definite	Established	None
Cnidarians-Hydrozoans	Gonionemus vertens	1894	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Algae	Gracilaria vermiculophylla	1998	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Crustaceans-Amphipods	Grandidierella japonica	2013	Definite	Established	Ballast Water, Hull Fouling
Algae	Grateloupia turuturu	1996	Definite	Established	Ballast Water, Hull Fouling
Platyhelminthes	Gyrodactylus anguillae*	1999	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster)
Protozoans	Haplosporidium nelsoni*	1957	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Annelids-Polychaetes	Hediste diversicolor	1963	Definite	Established	None
Crustaceans-Crabs	Hemigrapsus sanguineus	1988	Definite	Established	Ballast Water
Mollusks-Gastropods	Heterogen japonica	2000	Definite	Established	None
Bryozoans	Hippoporina indica	2001	Definite	Established	Hull Fouling
Arthropoda-Insects	Holcocranum saturejae*	1955	Definite	Established	None
Arthropoda-Insects	Holocranum saturejae*	1955	Definite	Established	None
Crustaceans-Amphipods	Hourstonius sp.	1994	Definite	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Hylobius transversovittatus*	1991	Definite	Established	None
Crustaceans-Isopods	Ianiropsis serricaudis	1999	Definite	Established	Ballast Water, Hull Fouling
Fishes	Ichthyomyzon unicuspis*	1974	Definite	Established	None
Fishes	Ictalurus furcatus	1974	Definite	Established	None
Fishes	Ictalurus punctatus	1889	Definite	Established	None
Mollusks-Bivalves	Ischadium recurvum	1869	Definite	Established	Oyster Accidental
Annelids-Polychaetes	Janua heterostropha	2000	Definite	Established	Hull Fouling
Fishes	Labidesthes sicculus*	1991	Definite	Established	None
Arthropoda-Insects	Lasioptera hungarica*	2000	Definite	Established	None
Fishes	Lepomis cyanellus	1900	Definite	Established	None
Fishes	Lepomis gulosus	1897	Definite	Established	None
Fishes	Lepomis macrochirus	1855	Definite	Established	None
Fishes	Lepomis megalotis*	1923	Definite	Established	None
Fishes	Lepomis microlophus	1974	Definite	Established	None
Crustaceans-Isopods	Ligia exotica	1924	Definite	Established	Cargo, Dry Ballast
Arthropoda-Insects	Lipara pullitarsis*	2002	Definite	Established	None
Arthropoda-Insects	Lipara rufitarsis*	2000	Definite	Established	None
Arthropoda-Insects	Lipara similis*	1998	Definite	Established	None
Mollusks-Gastropods	Littorina littorea	1875	Definite	Established	Dry Ballast, Hull Fouling, Natural Dispersal
Algae	Lomentaria clavellosa*	1972	Definite	Established	Ballast Water, Hull Fouling
Algae	Lomentaria orcadensis*	1853	Definite	Established	Hull Fouling
Bryozoans	Lophopodella carteri	1972	Definite	Established	None
Entoprocts	Loxosomatoides laevis	1995	Definite	Established	Hull Fouling
Crustaceans-Barnacles	Loxothylacus panopaei	1964	Definite	Established	Oyster Accidental
Crustaceans-Shrimp	Lysmata vittata	2013	Definite	Established	Ballast Water
Cnidarians-Hydrozoans	Maeotias marginata	1968	Definite	Established	Ballast Water, Hull Fouling
Algae	Melanosiphon intestinalis*	2002	Definite	Established	Ballast Water, Hull Fouling
Algae	Melanothamnus japonicus*	2007	Definite	Established	Ballast Water, Hull Fouling
Bryozoans	Membranipora membranacea	1990	Definite	Established	Ballast Water, Hull Fouling, Natural Dispersal
Crustaceans-Amphipods	Microdeutopus gryllotalpa	1872	Definite	Established	Hull Fouling
Fishes	Micropterus dolomieu	1855	Definite	Established	None
Fishes	Micropterus punctulatus	2005	Definite	Established	None
Fishes	Micropterus salmoides	1850	Definite	Established	None
Fishes	Misgurnus anguillicaudatus*	2013	Definite	Established	None
Cnidarians-Hydrozoans	Moerisia lyonsi	1965	Definite	Established	Ballast Water, Hull Fouling
Fishes	Morone chrysops*	1976	Definite	Established	None
Fishes	Moxostoma macrolepidotum*	1999	Definite	Established	None
Mollusks-Gastropods	Myosotella myosotis	1832	Definite	Established	Cargo, Dry Ballast
Mollusks-Bivalves	Mytilopsis leucophaeata	1937	Definite	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Nacerdes melanura*	1885	Definite	Established	None
Annelids-Polychaetes	Neodexiospira brasiliensis	1982	Definite	Established	Hull Fouling
Fishes	Neogobius melanostomus*	2021	Definite	Established	None
Algae	Neopyropia katadae*	2005	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster), Hull Fouling
Algae	Neopyropia koreana*	2003	Definite	Established	Ballast Water, Hull Fouling
Algae	Neopyropia yezoensis*	2005	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster), Hull Fouling
Fishes	Notropis atherinoides*	1936	Definite	Established	None
Mollusks-Bivalves	Ostrea edulis	1949	Definite	Established	Ballast Water, Natural Dispersal, Oyster Intentional
Crustaceans-Shrimp	Palaemon elegans	2017	Definite	Established	Ballast Water, Discarded Bait, Natural Dispersal
Crustaceans-Shrimp	Palaemon macrodactylus	2001	Definite	Established	Ballast Water
Fishes	Percina peltata*	1936	Definite	Established	None
Protozoans	Perkinsus marinus*	1949	Definite	Established	Oyster Accidental
Arthropoda-Insects	Petrobius brevistylis*	1979	Definite	Established	Cargo, Dry Ballast
Algae	Phycocalidia suborbiculata*	1966	Definite	Established	Ballast Water, Hull Fouling
Fishes	Pimephales promelas	1936	Definite	Established	None
Mollusks-Bivalves	Pisidium amnicum	1962	Definite	Established	None
Mollusks-Gastropods	Pleurocera canaliculata	1869	Definite	Established	None
Fishes	Pomoxis annularis	1894	Definite	Established	None
Fishes	Pomoxis nigromaculatus	1873	Definite	Established	None
Mollusks-Gastropods	Potamopyrgus antipodarum	2021	Definite	Established	None
Annelids-Oligochaetes	Potamothrix vejdovskyi*	1974	Definite	Established	None
Crustaceans-Mysids	Praunus flexuosus	2010	Definite	Established	Ballast Water, Natural Dispersal
Crustaceans-Crayfish	Procambarus clarkii	1981	Definite	Established	None
Crustaceans-Crayfish	Procambarus zonangulus	2006	Definite	Established	None
Arthropoda-Insects	Procanace dianneae*	1987	Definite	Established	Cargo, Hull Fouling
Platyhelminthes	Pseudodactylogyrus anguillae*	1999	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster)
Fishes	Pylodictis olivaris	1965	Definite	Established	None
Mollusks-Bivalves	Rangia cuneata	1957	Definite	Established	Ballast Water, Canal, Oyster Accidental
Mollusks-Gastropods	Rapana venosa	1998	Definite	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Rhizedra lutosa*	1988	Definite	Established	None
Algae	Rhodymenia delicatula*	1996	Definite	Established	Ballast Water, Hull Fouling
Fishes	Salmo trutta*	1885	Definite	Established	Fisheries Intentional
Fishes	Sander vitreus*	1879	Definite	Established	None
Fishes	Scardinius erythrophthalmus*	1986	Definite	Established	None
Arthropoda-Insects	Sclerocona acutella*	1984	Definite	Established	None
Crustaceans-Amphipods	Stenothoe gallensis species complex	1960	Definite	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Stomoxys calcitrans*	1781	Definite	Established	None
Mollusks-Gastropods	Stramonita canaliculata	1955	Definite	Established	Hull Fouling, Oyster Accidental
Algae	Striaria attenuata*	1848	Definite	Established	Hull Fouling
Tunicates	Styela canopus	1871	Definite	Established	Hull Fouling
Tunicates	Styela clava	1973	Definite	Established	Hull Fouling
Tunicates	Styela plicata	2002	Definite	Established	Hull Fouling
Crustaceans-Isopods	Synidotea laticauda	1999	Definite	Established	Ballast Water, Hull Fouling
Arthropoda-Insects	Telmatogeton japonicus*	1947	Definite	Established	Cargo, Hull Fouling
Mollusks-Bivalves	Teredo navalis	1862	Definite	Established	Hull Fouling
Arthropoda-Insects	Tetramesa phragmitis*	2000	Definite	Established	None
Arthropoda-Insects	Thambemyia borealis*	2015	Definite	Established	None
Mollusks-Gastropods	Thecacera pennigera	2003	Definite	Established	Ballast Water, Hull Fouling
Bryozoans	Tricellaria inopinata	2010	Definite	Established	Hull Fouling
Algae	Ulonema rhizophorum*	2002	Definite	Established	None
Algae	Ulva australis	2011	Definite	Established	None
Algae	Ulva laetevirens*	2011	Definite	Established	Ballast Water, Hull Fouling
Fishes	Umbra limi*	1974	Definite	Established	None
Mollusks-Gastropods	Valvata piscinalis	1985	Definite	Established	None
Bryozoans	Victorella pavida	1920	Definite	Established	Hull Fouling
Mollusks-Gastropods	Viviparus georgianus	1895	Definite	Established	None

* = Species summary still under review.

na-et3 (Cape Cod to Cape Hatteras) Summary