Taxonomic Group	Name	Year	Invasion Status	Population Status	Vectors
Fishes	Acanthogobius flavimanus	2023	Definite	Established	Natural Dispersal
Arthropoda-Insects	Aedes togoi*	None	Definite	Established	None
Annelids-Polychaetes	Alitta succinea	1976	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Fishes	Alosa sapidissima	1880	Definite	Established	Natural Dispersal
Fishes	Ameiurus catus	1930	Definite	Established	None
Fishes	Ameiurus melas	1894	Definite	Established	None
Fishes	Ameiurus natalis	1905	Definite	Established	None
Fishes	Ameiurus nebulosus	1894	Definite	Established	None
Crustaceans-Barnacles	Amphibalanus improvisus	1955	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Crustaceans-Amphipods	Ampithoe valida	1950	Definite	Established	Ballast Water, Discarded Bait, Hull Fouling, Oyster Accidental
Crustaceans-Amphipods	Aoroides secunda	1996	Definite	Established	Ballast Water, Hull Fouling
Mollusks-Gastropods	Assiminea parasitologica	2007	Definite	Established	Ballast Water, Cargo, Hull Fouling
Platyhelminthes	Austrobilharzia variglandis*	2003	Definite	Established	Oyster Accidental
Entoprocts	Barentsia benedeni	1988	Definite	Established	Hull Fouling, Oyster Accidental
Mollusks-Gastropods	Batillaria attramentaria	2007	Definite	Established	Fisheries Accidental (not Oyster), Oyster Accidental
Cnidarians-Hydrozoans	Blackfordia virginica	1997	Definite	Established	Ballast Water, Hull Fouling
Crustaceans-Cladocerans	Bosmina coregoni	2007	Definite	Established	None
Tunicates	Botrylloides violaceus	1979	Definite	Established	Hull Fouling, Oyster Accidental
Tunicates	Botryllus schlosseri	1972	Definite	Established	Hull Fouling, Oyster Accidental
Bryozoans	Bugula neritina	1986	Definite	Established	Hull Fouling, Oyster Accidental
Bryozoans	Bugulina stolonifera	2003	Definite	Established	Hull Fouling
Crustaceans-Isopods	Caecidotea racovitzai	1999	Definite	Established	None
Algae	Campylaephora kondoi*	1998	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Algae	Campylaephora sungminbooi*	1998	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Crustaceans-Amphipods	Caprella drepanochir	1988	Definite	Established	Ballast Water, Hull Fouling
Crustaceans-Amphipods	Caprella mutica	1973	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Fishes	Carassius auratus	1933	Definite	Established	None
Crustaceans-Crabs	Carcinus maenas	1995	Definite	Established	Ballast Water, Hull Fouling, Natural Dispersal
Mollusks-Gastropods	Catriona rickettsi	1994	Definite	Established	Ballast Water, Hull Fouling
Mollusks-Gastropods	Cecina manchurica	1963	Definite	Established	Oyster Accidental
Platyhelminthes	Cercaria batillariae*	2016	Definite	Established	Fisheries Accidental (not Oyster)
Sponges	Chalinula loosanoffi*	2001	Definite	Established	Hull Fouling, Oyster Accidental
Crustaceans-Amphipods	Chelura terebrans	1960	Definite	Established	Hull Fouling
Arthropoda-Insects	Chilacis typhae*	1998	Definite	Established	None
Tunicates	Ciona savignyi	2001	Definite	Established	Ballast Water, Hull Fouling
Mollusks-Gastropods	Cipangopaludina chinensis	2002	Definite	Established	None
Sponges	Clathria prolifera*	1967	Definite	Established	Hull Fouling, Oyster Accidental
Cnidarians-Hydrozoans	Clava multicornis	2013	Definite	Established	Discarded Bait, Hull Fouling
Cnidarians-Hydrozoans	Climacocodon ikarii	2002	Definite	Established	Ballast Water
Bryozoans	Conopeum chesapeakensis	2015	Definite	Established	Ballast Water, Hull Fouling
Bryozoans	Conopeum tenuissimum	1970	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Mollusks-Bivalves	Corbicula fluminea	1938	Definite	Established	None
Cnidarians-Hydrozoans	Cordylophora caspia	1959	Definite	Established	Aquatic Plant Shipments, Fisheries Accidental (not Oyster), Hull Fouling, Oyster Accidental
Tunicates	Corella inflata	2003	Definite	Established	Hull Fouling
Crustaceans-Amphipods	Crangonyx pseudogracilis	2002	Definite	Established	None
Mollusks-Gastropods	Crepidula convexa	1989	Definite	Established	Hull Fouling, Oyster Accidental
Mollusks-Gastropods	Crepidula fornicata	1937	Definite	Established	Oyster Accidental
Mollusks-Gastropods	Crepidula plana	1937	Definite	Established	Oyster Accidental
Bryozoans	Cryptosula pallasiana	1972	Definite	Established	Hull Fouling, Oyster Accidental
Fishes	Cyprinus carpio	1883	Definite	Established	None
Cnidarians-Anthozoans	Diadumene leucolena	1967	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Cnidarians-Anthozoans	Diadumene lineata	1947	Definite	Established	Hull Fouling, Oyster Accidental
Cnidarians-Anthozoans	Diadumene paranaensis	1988	Definite	Established	Ballast Water, Hull Fouling
Tunicates	Didemnum vexillum	2001	Definite	Established	Fisheries Accidental (not Oyster), Hull Fouling, Oyster Accidental
Annelids-Polychaetes	Diplocirrus SD1	2012	Definite	Established	None
Tunicates	Diplosoma listerianum	1988	Definite	Established	Hull Fouling, Oyster Accidental
Cnidarians-Hydrozoans	Ectopleura crocea	1948	Definite	Established	Hull Fouling
Annelids-Oligochaetes	Eukerria saltensis*	2003	Definite	Established	None
Crustaceans-Copepods	Eurytemora carolleeae	1998	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster), Oyster Accidental
Crustaceans-Ostracods	Eusarsiella zostericola	2000	Definite	Established	Oyster Accidental
Bryozoans	Fredericella indica	1999	Definite	Established	None
Algae	Fucus spiralis	1994	Definite	Established	Discarded Bait, Dry Ballast, Hull Fouling
Fishes	Fundulus diaphanus*	1971	Definite	Established	None
Arthropoda-Insects	Galerucella calmariensis*	1997	Definite	Established	None
Arthropoda-Insects	Galerucella pusilla*	1997	Definite	Established	None
Fishes	Gambusia affinis	1995	Definite	Established	None
Mollusks-Bivalves	Gemma gemma	1992	Definite	Established	Oyster Accidental
Algae	Gracilaria vermiculophylla	2017	Definite	Established	Oyster Accidental
Crustaceans-Amphipods	Grandidierella japonica	1977	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Sponges	Halichondria bowerbanki*	1977	Definite	Established	Hull Fouling, Oyster Accidental
Crustaceans-Copepods	Harpacticella paradoxa	2000	Definite	Established	None
Annelids-Polychaetes	Hediste diadroma	2009	Definite	Established	Ballast Water, Hull Fouling
Annelids-Polychaetes	Heteromastus filiformis species complex	1970	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Platyhelminthes	Himasthla quissetensis*	2003	Definite	Established	Oyster Accidental
Annelids-Polychaetes	Hobsonia florida	1980	Definite	Established	Hull Fouling, Oyster Accidental
Arthropoda-Insects	Hylobius transversovittatus*	1997	Definite	Established	None
Fishes	Hypomesus nipponensis	1959	Definite	Established	None
Crustaceans-Isopods	Iais californica	1931	Definite	Established	Ballast Water, Hull Fouling
Crustaceans-Isopods	Ianiropsis serricaudis	2015	Definite	Established	None
Fishes	Ictalurus punctatus	1945	Definite	Established	None
Mollusks-Gastropods	Ilyanassa obsoleta	1945	Definite	Established	Oyster Accidental
Crustaceans-Amphipods	Incisocalliope derzhavini	1986	Definite	Established	Ballast Water, Hull Fouling
Crustaceans-Amphipods	Jassa marmorata	1949	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Protozoans	Lagenophrys cochinensis*	1931	Definite	Established	Hull Fouling
Mollusks-Bivalves	Laternula gracilis	1995	Definite	Established	Ballast Water, Oyster Accidental
Fishes	Lepomis gibbosus	1893	Definite	Established	None
Fishes	Lepomis gulosus	1930	Definite	Established	None
Fishes	Lepomis macrochirus	1933	Definite	Established	None
Crustaceans-Copepods	Lernaea cyprinacea	1948	Definite	Established	None
Mollusks-Bivalves	Limecola petalum	1996	Definite	Established	Ballast Water, Oyster Accidental
Crustaceans-Copepods	Limnoithona tetraspina	2003	Definite	Established	Ballast Water
Crustaceans-Isopods	Limnoria quadripunctata	1949	Definite	Established	Hull Fouling
Crustaceans-Isopods	Limnoria tripunctata	1964	Definite	Established	Hull Fouling, Oyster Accidental
Algae	Lomentaria hakodatensis*	1990	Definite	Established	Hull Fouling, Oyster Accidental
Fishes	Lucania parva	1958	Definite	Established	Fisheries Accidental (not Oyster)
Mollusks-Bivalves	Magallana gigas	1928	Definite	Established	Oyster Intentional
Annelids-Polychaetes	Marenzelleria neglecta	2020	Definite	Established	Ballast Water
Annelids-Polychaetes	Megasyllis nipponica	2015	Definite	Established	None
Crustaceans-Amphipods	Melita nitida	1954	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Crustaceans-Amphipods	Microdeutopus gryllotalpa	1985	Definite	Established	Ballast Water, Hull Fouling
Fishes	Micropterus dolomieu	1924	Definite	Established	None
Fishes	Micropterus salmoides	1888	Definite	Established	None
Fishes	Misgurnus anguillicaudatus*	1997	Definite	Established	None
Tunicates	Molgula citrina	2010	Definite	Established	Hull Fouling
Tunicates	Molgula manhattensis	1974	Definite	Established	Hull Fouling, Oyster Accidental
Crustaceans-Amphipods	Monocorophium acherusicum	1905	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Crustaceans-Amphipods	Monocorophium insidiosum	1987	Definite	Established	Ballast Water, Hull Fouling
Crustaceans-Amphipods	Monocorophium uenoi	2000	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Fishes	Morone saxatilis	1887	Definite	Established	Fisheries Intentional, Natural Dispersal
Mollusks-Bivalves	Mya arenaria	1875	Definite	Established	Fisheries Intentional, Oyster Accidental
Mollusks-Gastropods	Myosotella myosotis	1876	Definite	Established	Cargo, Dry Ballast, Oyster Accidental
Crustaceans-Copepods	Mytilicola orientalis	1938	Definite	Established	Hull Fouling, Oyster Accidental
Mollusks-Bivalves	Mytilus galloprovincialis	1996	Definite	Established	Ballast Water, Fisheries Accidental (not Oyster), Hull Fouling
Arthropoda-Insects	Nanophyes marmoratus*	1997	Definite	Established	None
Cnidarians-Anthozoans	Nematostella vectensis	1973	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Mollusks-Bivalves	Neotrapezium liratum	1947	Definite	Established	Oyster Accidental
Annelids-Polychaetes	Nicolea sp. A. Harris	2011	Definite	Established	Ballast Water, Hull Fouling
Crustaceans-Cumaceans	Nippoleucon hinumensis	1977	Definite	Established	Ballast Water, Oyster Accidental
Fishes	Notemigonus crysoleucas	2002	Definite	Established	None
Mollusks-Bivalves	Nuttallia obscurata	2002	Definite	Established	Ballast Water, Natural Dispersal
Mollusks-Gastropods	Ocinebrellus inornatus	1934	Definite	Established	Oyster Accidental
Crustaceans-Isopods	Orthione griffenis	1988	Definite	Established	Ballast Water
Crustaceans-Crayfish	Pacifastacus leniusculus	1959	Definite	Established	None
Crustaceans-Shrimp	Palaemon macrodactylus	1987	Definite	Established	Ballast Water
Crustaceans-Shrimp	Palaemon modestus	1995	Definite	Established	None
Crustaceans-Amphipods	Paracorophium sp.	2000	Definite	Established	Ballast Water, Hull Fouling
Bryozoans	Pectinatella magnifica	1998	Definite	Established	None
Fishes	Perca flavescens	1891	Definite	Established	None
Tunicates	Perophora japonica	2003	Definite	Established	Hull Fouling
Mollusks-Bivalves	Petricolaria pholadiformis	1943	Definite	Established	Oyster Accidental
Mollusks-Gastropods	Philine auriformis	1998	Definite	Established	Ballast Water, Natural Dispersal
Mollusks-Gastropods	Philine orientalis	2015	Definite	Established	Ballast Water
Annelids-Polychaetes	Polydora cornuta	1950	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Fishes	Pomoxis annularis	1893	Definite	Established	None
Fishes	Pomoxis nigromaculatus	1893	Definite	Established	None
Mollusks-Gastropods	Potamopyrgus antipodarum	1997	Definite	Established	None
Arthropoda-Insects	Prokelisia marginata*	2000	Definite	Established	Biocontrol
Crustaceans-Copepods	Pseudodiaptomus forbesi	1999	Definite	Established	Ballast Water
Crustaceans-Copepods	Pseudodiaptomus inopinus	1990	Definite	Established	Ballast Water
Annelids-Polychaetes	Pseudopolydora cf. kempi	1974	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Annelids-Polychaetes	Pseudopolydora paucibranchiata	1976	Definite	Established	Ballast Water, Oyster Accidental
Crustaceans-Isopods	Pseudosphaeroma sp. A	2003	Definite	Established	Ballast Water, Hull Fouling
Fishes	Ptychocheilus grandis	1989	Definite	Established	None
Fishes	Rhinogobius brunneus species complex*	2004	Definite	Established	Ballast Water
Crustaceans-Crabs	Rhithropanopeus harrisii	1950	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Mollusks-Bivalves	Ruditapes philippinarum	1947	Definite	Established	Fisheries Intentional, Oyster Accidental
Fishes	Sander vitreus*	1981	Definite	Established	None
Algae	Sargassum muticum	1947	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Bryozoans	Schizoporella japonica	1970	Definite	Established	Hull Fouling, Oyster Accidental
Bryozoans	Schizoporella variabilis	2015	Definite	Established	None
Crustaceans-Tanaids	Sinelobus cf. stanfordi	1989	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Crustaceans-Copepods	Sinocalanus doerrii	1999	Definite	Established	None
Crustaceans-Isopods	Sphaeroma quoianum	1931	Definite	Established	Ballast Water, Hull Fouling
Crustaceans-Amphipods	Stenothoe valida	2000	Definite	Established	Ballast Water, Hull Fouling
Annelids-Polychaetes	Streblospio benedicti	1976	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Tunicates	Styela clava	1993	Definite	Established	Hull Fouling
Annelids-Oligochaetes	Stylodrilus heringianus*	2001	Definite	Established	None
Crustaceans-Isopods	Synidotea laticauda	1989	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Mollusks-Gastropods	Tenellia adspersa	1986	Definite	Established	Ballast Water, Hull Fouling
Mollusks-Bivalves	Teredo navalis	1957	Definite	Established	Hull Fouling
Protozoans	Trochammina hadai*	1998	Definite	Established	Ballast Water, Oyster Accidental, Unknown Vector
Annelids-Oligochaetes	Tubificoides brownae*	1986	Definite	Established	Ballast Water, Dry Ballast, Oyster Intentional
Annelids-Oligochaetes	Tubificoides diazi*	1979	Definite	Established	Ballast Water, Dry Ballast, Hull Fouling, Oyster Accidental
Algae	Ulva australis	2010	Definite	Established	Ballast Water, Hull Fouling, Oyster Accidental
Entoprocts	Urnatella gracilis	2002	Definite	Established	None
Mollusks-Gastropods	Urosalpinx cinerea	1941	Definite	Established	Oyster Accidental
Bryozoans	Watersipora n. sp.	2007	Definite	Established	Hull Fouling

* = Species summary still under review.

nep-iv (Puget Sound to Northern California) Summary